The Cat Survival Trust


The Clouded Leopard

Neofelis nebulosa Raffles or Griffith
A clouded leopard in the wild

Contents

 
 
  • Description
  • Distribution
  • Diet
  • Behaviour
  • Reproduction
  • Conservation Status
  • Captive Breeding and Clouded Leopards in Captivity
  • Further Reading

  • A Clouded Leopard in the Indonesian Forest
    Photo: Robert C. Martin


    Other names

      
     French:Panthère longibande, panthère nébuleuse
     German:Nebelparder
     Spanish:pantera longibanda, pantera nebulosa

    Description

    Sir Stamford Raffles was the first person to describe the clouded leopard, but Griffith was the first to ascribe a scientific name to it. The Malaysians call this animal the “Rimau-dahan” or “Harimau-dahan” which translate as Tree or Branch tiger. It is very elusive and has rarely been seen.
    The clouded leopard has been described as having the skull and dentition of a big cat with the body of a small cat. It is the size of a small leopard, with a long body and tail and short limbs. The coat is earthy-brown, pale or a rich yellowish brown. Melanistic or all black individuals are thought to exist. Its name describes the large blotchy cloudlike markings on its body. They have a dark margin and a pale centre. Dark spots cover the head, tail and limbs, and big black spots mark the white or pale tawny underparts. On the cheeks and sides of the head are two black bands. One runs from the eye to beneath the ear, the other is parallel, running back from the mouth. The ears themselves are relatively short and rounded, their backs are black with a greyish central patch.
    The tail is incompletely ringed and is almost as long as the body. Long tails are used as counterbalances when climbing.
    In comparison to the marbled cat, which it resembles, the skull is long and narrow. The face of the clouded leopard appears to be more like that of a big cat, whereas the marbled cat has a short rounded face. Like the small cats, the hyoid bones which support the clouded leopard’s larynx are bony, this in conjunction with flexible vocal cords, means that they can purr continuously.
    The ankle joints of the clouded leopard are notably flexible, although not as much as the margay’s. This is a substantial aid to climbing. Wide strong paws enable the clouded leopard to run along the underside of a branch. They are said to be able to hang by one hind foot from a branch as they lie in ambush.
    The eyes of clouded leopards are distinctive in that the pupils are vertically oblong rather than round or linear as in other cats.
    The upper canines of the clouded leopard are relatively much longer than any other feline, almost tusk-like. Canines 4.4 cm long have been recorded. A broad gap between the canines and premolars (the anterior upper premolar is absent) enables the clouded leopard to take very big bites. The posterior edge is very sharp, like the canines of the prehistoric sabre-toothed cats. Due to these unique characteristics the clouded leopard has been allocated a genus to itself.
    Four subspecies have been described although as with all subdivisions of species they are not universally recognised:
    N. n. nebulosaSouth China, Indo-China
    N. n. brachyurusTaiwan (Extinct?)
    N. n. diardiBorneo
    N. n. macrosceloidesNepal to Burma

    Principal dimensions

     OverallMalesFemales
    Head and body lengths (cm)60-110  
    Height at shoulder (cm)53  
    Tail lengths (cm)55-91  
    Weight (Kg)15-30  

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    Distribution and Habitats

    The clouded leopard is an animal of the tropical forests, up to altitudes of about 2,500 metres. It seems to be able to survive in marginal scrub and in selectively logged secondary forest. It is found throughout south east Asia, from Nepal, Bhutan and southern China to Malaysia, Borneo and Sumatra. Bangjie (1984) stated that the clouded leopard has a wider distribution in China than had been thought. He reported that it could be found as far north as the north west border. It was endemic to Taiwan but has probably been hunted to extinction.
    The map shows the present distribution of Clouded Leopards in grey.
    The map is based on information in the Wild Cats Status Survey and Conservation Action Plan published by the IUCN/SSC Cat Specialist Group in 1996. See our
    Books page for more details.
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    Diet

    Clouded leopards are opportunistic predators, as are most cats. Recorded prey are sambar, muntjac, birds, bearded pig, civet, monkeys, gibbons, squirrels, porcupines, fish, and domestic cattle and chickens. Captive animals will eat eggs and some vegetation.
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    Behaviour


    Photo: Terry Moore

    Apparently highly adapted for an arboreal existence, the Bornean clouded leopards are said to be mostly terrestrial, those elsewhere live more or less in the trees. Panwar and Gopal (1984) state than due to their short legs they are rather clumsy when on the ground. Their behaviour is flexible and they use roads and artificial trails to travel and hunt.
    They have been seen during the day, but their activity periods are mainly crepuscular. Resting individuals have been observed in nests, high in trees. Rabinowitz et al (1987) report that they sleep on top of boulders in rocky areas and suggest that this behaviour could be an adaptation to avoid being eaten by ground-dwelling leeches.
    When eating, the clouded leopard’s posture is close to that of the large or Panthera cats; they lie down to feed, rather than crouch as small cats do.
    The social systems of clouded leopards are unknown but it has been reported that a couple may stay together for a period. Certainly in captivity, some males have been known to tolerate the female and her kittens and vice versa although, sadly there have been more reports of fatal fights between captive cats kept in the same enclosure.
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    Reproduction

    Most litters consist of two cubs but four or even five have been born in captivity. Gestation periods range from 86 to 109 days.
    The cubs weigh about 140 to 280 grammes at birth and are weaned in 93 days. Their eyes are open by 12 days and the cubs can walk when they are about 20 days old. If allowed, they will continue to suckle for five months.
    Males are said to develop faster than females and they achieve adult coloration when they are about six months old.
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    Conservation Status

    In India the clouded leopard seems to have benefited from the efforts to conserve the tiger (Panwar and Gopal 1984). Both animals share the same habitat, so both gain from the tiger reserves. However these cats provide extremely desirable products. Markets in Bangladesh have been flooded with clouded leopard skins (Khan 1984). The remaining Bangladeshi populations are sustained and replenished by individuals from the neighbouring forests in India and Burma.
    Rabinowitz et al (1987) state that the clouded leopard, the largest felid on Borneo, is still present in some numbers. It is not frequently encountered or specifically hunted and there is virtually no market for skins or bodyparts. Although the natives often shoot it on sight it does not seem to be in imminent danger of extinction.
    Although the clouded leopard is protected in China, Bangjie (1984) reports that pelts are openly bought and sold. 119 were declared caught in 1980/81. He implies that there is a substantial breeding population in Chinese zoos and concludes that clouded leopards are rare, but not as much so as the larger cats.
    Killing a clouded leopard bestowed heroic status on the hunter in Taiwan (Nowell 1990b). Hunting them has been illegal for the past 20 years but their skins remain a great badge of wealth. There is now a lucrative market for Chinese smugglers. Nowell (1990a) states that the most recent sighting of a wild Formosan clouded leopard was by a biologist in 1983, the animal was a snared juvenile. She reports that there is a large amount of protected habitat, high in the mountains. The clouded leopard’s historical distribution is below 2,500 metres, Nowell hopes that it can adapt to live higher. As far as is known, there are no representatives of this subspecies in captivity.
    Habitat destruction and human persecution have placed the clouded leopard in a ‘Vulnerable' classification according to the IUCN (Conservation and Legal Status of Wild Cats, Cat News 12, 1990, p.26).
    The Convention on International Trade in Endangered Species (CITES) has placed it on Appendix I, prohibiting all international commerce.
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    Captive Breeding and Clouded Leopardsin Captivity

    Studbook Keeper
    Norah B. Fletchall
    Assistant Director
    John Ball Zoological Gardens
    1300 W. Fulton Street NW
    Grand Rapids
    Michigan 49504, USA

    A captive breeding programme needs to have regard to keeping separate the various subspecies, while at the same time avoiding the dangers of inbreeding.
    This is achieved by the maintenance of a studbook which is used to select suitable breeding partners for captive animals. Such studbooks are maintained on a voluntary basis by dedicated individuals or teams at various zoos around the world.

    Zoos with Clouded Leopards


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    Last revision 11th June, 2002


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